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Grape-ferns are seedless vascular plants of the genus Botrypus, closely allied to (and previously often included in) the genus Botrychium (moonworts). They are small, with fleshy roots, and reproduce by spores shed into the air. They differ from the moonworts in having at least some sterile fronds (all fronds in Botrychium are spore-bearing), and in the fronds being bi- or tri-pinnate (Botrychium are single pinnate, or rarely bipinnate).
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