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The mantle is entirely within the shell. The foot extends from the larger end of the shell, and is used to burrow through the substrate. A number of minute tentacles around the foot, called captacula, sift through the sediment and latch onto bits of food which they then convey to the mouth. The mouth has grinding teeth that break the bit into smaller pieces for digestion. The scaphopid vascular system is rudimentary, lacking both heart and blood vessels; the blood is held in sinuses throughout the body cavity, and pumped by the rhythmic action of the foot. Tusk shells are well-known in the fossil record, first appearing in the Ordovician (last of all molluscan groups). The shells were used by the natives of the Pacific Northwest as wampum. References
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